The Genetic Theory Of Adaptation : A Brief History By H. Allen Orr

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In The Genetic Theory of Adaptation: A Brief History by H. Allen Orr the article discusses adaptation and how it was affected by micromutationism, phenotypic evolution, and sequence evolution and various models that arose through the years. Prior to getting into detail the article explains that “adaptation is not natural selection,” as emphasized in the 1930s by Ronald A. Fisher (Orr 1). It glosses over how evolutionary geneticists define and distinguish between the two terms and how the recent models on adaptation characterize evolutional patterns and fitness.
Micromutationism was a pre-Mendelian theory that began with Darwin and emphasized the gradual progression of phenotypic evolution. This theory was incredibly influential and had Biometric Schools founded upon its idea, however, it was ardently challenged by the then rising Mendelian genetics. Despite conflict the Micromutationism theory did eventually win out in the 1930s due mostly to Fisher and his blending of the two theories in order to construct “a mathematical framework known as the infinitesimal model,” (Orr 2). Though modern evolutionists tend to disregard this model it does have some credible points. The article then goes on to discuss two new theories that arose known as the quantitative trait locus analysis and the microbial experimental evolution. These theories concluded that evolution involved large genetic changes that came about through a compilation of a minimal number of minute changes.
In the next part of the article Fisher’s model on geometric adaptation is presented. This model presents small step evolution or gradualism. It conveys the effect that spontaneous mutations have on various traits. Fisher argued through his model that more complex spe...

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...for more accurate predictions of how species will evolve and whether or not certain species will survive and what adaptations could promote or inhibit that process. There are different methods of adaptation available for animals and plants to employ in nature. Thus, exploring adaptations during our lectures would grant us better insight into what caused certain things to demonstrate specific adaptations and would aid in the explanation and expansion of convergent evolution.
Understanding adaptation and reading about the processes and models developed to explain it allowed for the difference between adaptation and natural selection to cement themselves. Knowing that adaptation fits an organism to its environment, whereas natural selection eliminates those not fit to survive impacts an individual’s understanding of how evolution took place and continues to take place.

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