Succession

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Succession

Succession is where a pioneer species colonise an area, and gradually

bring about changes so that other more complex species may also

colonise the area, replacing the pioneers. The presence of the pioneer

species signifies primary succession, which is basically the

colonisation of bare rock. The abiotic conditions are extreme, with a

very harsh climate which can only be survived by the pioneer species.

An example of a pioneer species is lichen, which are found in the

splash zone of a shore. The abiotic conditions of the splash zone are

extremely harsh, with fluctuating temperature and prolonged exposure

to sunlight causing desiccation. Lichen is well adapted to survive

these conditions, as it's actually two organisms forming a mutualistic

symbiotic relationship. Fungus and algae live together, which is

beneficial to both organisms. The algae photosynthesises, which

provides carbohydrates for itself and the fungus. The fungus has

hyphae, which provides anchorage for the lichen as it can get into

tiny cracks in the rock. It also takes in water through osmosis, and

even minerals. The hyphae even protect the lichen from desiccation, as

it forms a mat which protects the lichen. Physical and chemical

weathering allow colonisation to begin. For example, when freeze/thaw

causes cracks to appear in the rocks, they provide ideal sheltered

areas for colonisation. Also, as rainwater is slightly acidic, wears

down the rocks, also causing suitable conditions.

When lichen die, the nitrogenous compounds are broken down by

saprophytic bacteria into ammonia or ammonium ions. Then, nitrifying

bacteria converts the ammonia compounds into nitrites, and then

quickly converted to nitrates due to the toxic nature of nitrites.

Nitrates are soluble in water and can be taken up by small plants such

as mosses by osmosis. After a few seral stages larger plants colonise,

which need to have deeprooting systems for anchorage. These plants can

therefore attract invertebrates and other organisms. Overall, the

index of diversity increases over time. Eventually, when the

environment is stable and the index of diversity is high, a climax

community has been formed.

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