desert tortoise

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Introduction
The desert tortoise is a relic symbol of the arid environment (Morafka & Berry 2002). But despite the robust array of evidence suggesting phylogeny, certain populations of the desert tortoise remain elusive (Berry et al. 2002). Genetic evaluation has resolved some phylogeny issues by establishing a new species within the clade (Murphy et al. 2011).
Traits and behaviors also play a role in the evolutionary history of the desert tortoise. Many traits and behaviors exhibited by the desert tortoise may be symplesiomorphies or exaptations rather than desert adaptations (Morafka & Berry 2002). Other characteristics of the desert tortoise include behavioral modification to drought and variation between sexes in space use (Duda et al. 1999; Harless et al. 2009).
Upper respiratory tract disease is also characteristic of the desert tortoise. Varying levels of Mycoplasma agassizii may cause clinical symptoms of the disease. An enzyme-linked immunosorbent assay (ELISA) was used to detect M. agassizii antibodies in tortoise blood. Two factors were concluded to affect the mortality and morbidity rate of URTD—climate and M. agassizii antibodies within tortoise blood (Sandmeier et al. 2009; Sandmeier et al. 2013).
Finally, conservation efforts such as translocation of desert tortoises into wild populations were assessed. Gene flow and genetic variability were described among different populations of desert tortoise (Murphy et al. 2007; Edwards et al. 2004; Edwards & Berry 2013).
The ‘Species’ Problem
Species are the fundamental unit of evolution and measurement for biodiversity conservation. Speciation occurs by the splitting of a subpopulation from its main population, resulting in phenotypic and genotypic change, both of which ...

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... these populations include the need for the placement of culverts and corridors (Edwards et al. 2004).
Conclusion
Due somewhat to the paucity of research, the desert tortoise remains an enigmatic species (Berry et al. 2002). This fact is troubling particularly because the species habitats have become so fragmented and encroached upon. Management strategies to conserve the desert tortoise require a multifaceted approach of genetic, behavioral and habitat assessment (Edwards et al. 2004). Further possible effects of climate change on the desert tortoise have not been assessed although it has been noted that changes in El Nino patterns may disrupt foraging plant material for juvenile desert tortoises (McCoy 2011). The patterns of ecological decline for the desert tortoise may react synergistically in unknown ways making extinction a very viable threat to these species.

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