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The purpose of the study was to determine if human modification has an effect on species co-occurrence in stream invertebrates using a combination three methods: experimentation, field survey, and analysis of null models. Researchers hypothesized that human habitat modification disrupts patterns of species co-occurrence. They also hypothesize that the observations will be consistent along spatial scales. It was assumed from prior research, completed by other researchers, that undisturbed communities would show negative species co-occurrence due to predation, competition, and habitat segregation. It was also assumed that disturbance to a habitat would weaken biotic interaction which is explained to lead to random associations between species. The background given efficiently explains everything that is needed to interpret the rest of the investigation.
Different scales were used to compare macroinvertebrate community assembly in natural and modified habitats. At the catchment scale, 32 reaches were studies. Seventeen drained pastured grassland and fifteen drained semi natural vegetation. Hand net collections were taken at 3 minute intervals. At the patch scale, communities in undisturbed substrate were compared with communities under attack by fine sediment due to upstream erosion. Eight streams and a total of sixty stream patches were sample using a surper-sampler. At the microhabitat scale, controlled in situ conditions were used to reproduce fine sediment deposition. Three streams with low loads of sediment were examined. Sixty five total trays full of natural substrate free of animals were used. They were placed at the predetermined sites. Twenty-one trays were used as control, 23 trays were treated with fin...

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...e not solid enough in this experiment to make any final ruling in either direction. The researchers point that most of the literature on this topic is comprised of looking for changes in species richness, abundance, and composition. They claim to have changed that with this research and they have shown that habitat modification can disrupt species co-occurrence but not relative abundance and changes in richness. They also point out that this contradicts current literature on the topic. There is still more research on the topic that needs to be completed. There values in species richness and relative abundance stayed close to one another but there were large variations seen at the highest sediment level. The p values were significant but as there is other literature that contradicts this point it is evident that more studies such as this need to be performed.

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